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Migrate Bird

Bird Migration

Bird migration is the regular seasonal journey undertaken by many species of birds. Bird movements include those made in response to changes in food availability, habitat or weather. These however are usually irregular or in only one direction and are termed variously as nomadism, invasions, dispersal or irruptions. Migration is marked by its annual seasonality.[1] In contrast, birds that are non-migratory are said to be resident or sedentary.

Bird Migration

General patterns

Many bird populations migrate long distances along a flyway. The most common pattern involves flying north in the spring to breed in the temperate or Arctic summer and returning in the fall to wintering grounds in warmer regions to the south.

The primary advantage of migration is conservation of energy. The longer days of the northern summer provide greater opportunities for breeding birds to feed their young. The extended daylight hours allow diurnal birds to produce larger clutches than those of related non-migratory species that remain in the tropics year round. As the days shorten in autumn, the birds return to warmer regions where the available food supply varies little with the season.

General patterns

These advantages offset the high stress, physical exertion costs, and other risks of the migration. Predation can be heightened during migration; the Eleonora's Falcon which breeds on Mediterranean islands has a very late breeding season, coordinated with the autumn passage of southbound passerine migrants, which it feeds to its young. A similar strategy is adopted by the Greater Noctule bat, which preys on nocturnal passerine migrants. The higher concentrations of migrating birds at stopover sites make them prone to parasites and pathogens, which require a heightened immune response.

Within a species not all populations may be migratory; this is known as "partial migration". Partial migration is very common in the southern continents; in Australia, 44% of non-passerine birds and 32% of passerine species are partially migratory. In some species, the population at higher latitudes tends to be migratory and will often winter at lower latitude. The migrating birds bypass the latitudes where other populations may be sedentary, where suitable wintering habitats may already be occupied. This is known as leap-frog migration. Within a population, there can also be different patterns of timing and migration based on the age groups and sex. Only the female Chaffinches in Scandinavia migrate, with the males staying resident. This has given rise to the latter's specific name of coelebs, a bachelor.

Most migrations begin with the birds starting off in a broad front. In some cases the migration may involve narrow belts of migration that are established as traditional routes termed as flyways. These routes typically follow mountain ranges or coastlines, and may take advantage of updrafts and other wind patterns or avoid geographical barriers such as large stretches of open water. The specific routes may be genetically programmed or learned to varying degrees. The routes taken on forward and return migration are often different.

Many of the larger birds fly in flocks. Flying in flocks helps in reducing the energy needed. Many large birds fly in a V-formation, which helps individuals save 12–20 % of the energy they would need to fly alone. Red Knots Calidris canutus and Dunlins Calidris alpina were found in radar studies to fly 5 km per hour faster in flocks than when they were flying alone.

The altitude at which birds fly during migration varies. An expedition to Mt. Everest found skeletons of Pintail and Black-tailed Godwit at 5000 m (16,400 ft) on the Khumbu Glacier. Bar-headed Geese have been seen flying over the highest peaks of the Himalayas above 8000 m (29000 ft) even when low passes of 3000 m (10000 ft) were nearby. Seabirds fly low over water but gain altitude when crossing land, and the reverse pattern is seen in landbirds.[12][13] However most bird migration is in the range of 150 m (500 ft) to 600 m (2000 ft). Bird-hit aviation records from the United States show most collisions occur below 600 m (2000 ft) and almost none above 1800 m (6000 ft).

In contrast, most species of penguin migrate by swimming. These routes can cover over 1000 km. Blue Grouse Dendragapus obscurus perform altitudinal migration mostly by walking. Emus in Australia have been observed to undertake long-distance movements on foot during droughts.
 

Historical views

The earliest recorded observations of bird migration were 3000 years ago, as noted by Hesiod, Homer, Herodotus, Aristotle and others. The Bible also notes migrations, as in the Book of Job (39:26), where the inquiry is made: "Doth the hawk fly by Thy wisdom and stretch her wings toward the south?" The author of Jeremiah (8:7) wrote: "The stork in the heavens knoweth her appointed time; and the turtledove, and the crane, and the swallow, observe the time of their coming."

Aristotle noted that cranes traveled from the steppes of Scythia to marshes at the headwaters of the Nile. Pliny the Elder, in his Historia Naturalis, repeats Aristotle's observations. Aristotle however suggested that swallows and other birds hibernated. This belief persisted as late as 1878, when Elliott Coues listed the titles of no less than 182 papers dealing with the hibernation of swallows. It was not until early in the nineteenth century that migration as an explanation for the winter disappearance of birds from northern climes was accepted.[15]

The discovery in Germany of white storks embedded with African arrows provided early clues on migration. One of the oldest of these Pfeilstorch specimens was found in 1822 near the German village of Klütz, in the state of Mecklenburg-Vorpommern.

Orientation and navigation

Orientation and navigation

Navigation is based on a variety of senses. Many birds have been shown to use a sun compass. Using the sun for direction involves the need for making compensation based on the time. Navigation has also been shown to be based on a combination of other abilities including the ability to detect magnetic fields (magnetoception), use visual landmarks as well as olfactory cues.

Long distance migrants are believed to disperse as young birds and form attachments to potential breeding sites and to favourite wintering sites. Once the site attachment is made they show high site-fidelity, visiting the same wintering sites year after year.

The ability of birds to navigate during migrations cannot be fully explained by endogenous programming, even with the

help of responses to environmental cues. The ability to successfully perform long-distance migrations can probably only be fully explained with an accounting for the cognitive ability of the birds to recognize habitats and form mental maps. Satellite tracking of day migrating raptors such as Ospreys and Honey Buzzards has shown that older individuals are better at making corrections for wind drift.

As the circannual patterns indicate, there is a strong genetic component to migration in terms of timing and route, but this may be modified by environmental influences. An interesting example where a change of migration route has occurred because of such a geographical barrier is the trend for some Blackcaps in central Europe to migrate west and winter in Britain rather than cross the Alps.

Migratory birds may use two electromagnetic tools to find their destinations: one that is entirely innate and another that relies on experience. A young bird on its first migration flies in the correct direction according to the Earth's magnetic field, but does not know how far the journey will be. It does this through a radical pair mechanism whereby chemical reactions in special photo pigments sensitive to long wavelengths are affected by the field. Note that although this only works during daylight hours, it does not use the position of the sun in any way. At this stage the bird is similar to a boy scout with a compass but no map, until it grows accustomed to the journey and can put its other facilities to use. With experience they learn various landmarks and this "mapping" is done by magnetites in the trigeminal system, which tell the bird how strong the field is. Because birds migrate between northern and southern regions, the magnetic field strengths at different latitudes let it interpret the radical pair mechanism more accurately and let it know when it has reached its destination. More recent research has found a neural connection between the eye and "Cluster N", the part of the forebrain that is active during migrational orientation, suggesting that birds may actually be able to see the magnetic field of the earth.

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